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  • Landscape Biodiversity Lab

    Montana State University
    310 Lewis Hall
    Bozeman, MT 59717


    Andrew J. Hansen

    Tel: (406) 994-6046
    E-mail: hansen at



    Hansen, A.J., Piekielek, N., Davis, C., Haas, J., Theobald, D., Gross, J., Monahan, W., Olliff, T., Running, S., 2014. Exposure of U.S. National Parks to land use and climate change 1900-2100, Ecological Applications, 24(3), pp. 484-502. .pdf

    Goetz, S.J. Sun, M., S. Zolkos, A. Hansen and R. Dubayah, 2014. The relative importance of climate and vegetation properties on patterns of North American breeding bird species richness, Environmental Research Letters, 9:034013. .pdf

    Stine, P., P. Hessburg, T. Spies, M. Kramer, C. Fettis, A. Hansen, J. Lehmkuhl, K. O'Hara, K. Polivka, P. Singleton, S. Charnley, and A. Merschel. 2013. The Ecology and management of Moist Mixed-conifer forests in Eastern Oregon and Washington: a synthesis of the relevant biophysical science and implications for future land management (DRAFT),

    Hansen, A.J., N. Piekielek, C. Davis, J. Haas, D. Theobald, J. Gross, W. Monahan, S. Running.  In press.  Exposure of US National Parks to Land Use and Climate Change 1900-2100.  Ecological Applications. . OR .pdf

    Powell, S.J., A.J. Hansen, T.J. Rodhouse, L.K. Garrett, J.L. Betancourt, et al. 2013. Woodland Dynamics at the Northern Range Periphery: A Challenge for Protected Area Management in a Changing World. PLoS ONE 8(7): e70454. doi:10.1371/journal.pone.0070454. PDF

    Piekielek, NB and AJ Hansen. 2012. Extent of fragmentation of coarse-scale habitats in and around US National Parks. Biological Conservation 155:13-22. PDF

    Piekielek, N.B. 2012. Remote sensing grassland phenology in the Greater Yellowstone Ecosystem: biophysical correlates, land use effects and patch dynamics.Ph.D. Dissertation. Montana State University, Bozeman, MT. PDF

    Hansen, A.J. 2011.  Land-use change in the Greater Yellowstone Ecosystem: Past, Present and Possible Future Patterns and Consequences, in Questioning Greater Yellowstone's Future Climate, Land Use, and Invasive Species, the 10th Biennial Scientific Conference on the Greater Yellowstone Ecosystem. PDF

    Davis, C.R. and A.J. Hansen. 2011.  Trajectories in land-use change around US National Parks and their challenges and opportunities for management.  Ecological Applications 21(8) 3299-3316. PDF

    Hansen, A.J., L.B. Phillips, C.H. Flather and J. Robison-Cox. 2011. Carrying capacity for species richness as a context for conservation: A case study of North American breeding birds.  Global Ecology and Biogeography 20, 817-831. PDF and Cover

    Baril, L.M., A.J. Hansen, R. Renkin, R. Lawrence. 2011.  Songbird response to increased willow (Salix spp.) growth in Yellowstone's northern range.  Ecological Applications 21(6) 2283-2296 PDF

    Hansen, A. J., C. Davis, N. B. Piekielek, J. Gross, D. Theobald, S. Goetz, F. Melton, R. DeFries.  2011.  Delineating the ecosystems containing protected areas for monitoring and management.  BioScience 61(5) 363-373. PDF

    Phillips, L.B., Hansen, A.J., C.H. Flather and J. Robison-Cox. 2010. Applying species-energy theory to conservation: a case study for North American birds. Ecological Applications 20(7) 2207-2023.    PDF

    Olliff, T., G. Plumb, J. Kershner, C. Whitlock, A. Hansen, M. Cross, S. Bischke. 2010. A science agenda for the Greater Yellowstone Area. Yellowstone Science 18(2):14-22. PDF

    Baril, L.M., A.J. Hansen, R. Renkin, R. Lawrence. 2009. Willow-bird relationships on the Northern Range. Yellowstone Science 17(3):19-26. PDF

    Hansen, A.J.  2009.  Species and habitats most at risk in Greater Yellowstone.  Yellowstone Science 17(3):27-36. PDF

    McWethy, D.B., A.J. Hansen, and J.P. Verschuyl. 2010. Bird response to disturbance varies with forest productivity in the northwestern United States, Landscape Ecology 25(4): 533-549.    PDF

    DeFries, R., F. Rovero, P. Wright, J. Ahumada, S. Andelman, K. Brandon, J. Dempewolf, A. Hansen, J. Hewson, J. Liu.  2010.  Linking plot-level biodiversity measurements with human influences over multiple spatial scales in the tropics: A conceptual framework.  Frontiers in Ecology and the Environment 8(3): 153-160. PDF

    Jones, D.A., A.J. Hansen, K. Bly, K. Doherty, J.P. Verschuyl, J.I. Paugh, R. Carle, and S.J. Story. 2009. Monitoring land use and cover around parks: a conceptual approach.  Remote Sensing of Environment 113:1346-1356.  PDF

    McWethy, D.B., A.J. Hansen, and J.P. Verschuyl. 2009. Edge effects for songbirds vary with forest productivity.  Forest Ecology and Management 257, 665-678.    PDF

    Phillips, L.B., Hansen, A.J., and C.H. Flather, 2008. Evaluating the species energy relationship with the newest measures of ecosystem energy: NDVI versus MODIS primary production, Remote Sensing of Environment 112:4381-4392.  PDF

    Verschuyl, J.P., A.J. Hansen, D.B. McWethy, R. Sallabanks, R.L. Hutto.  2008.  Is the effect of forest structure on bird diversity modified by forest productivity?  Ecological Applications 18(5), 1155-1170.     PDF

    Pearce, J.L., D.A Kirk, C.P Lane, M.H. Mahr, J.Walmsley, D. Casey, J. Muir, S. Hannon, A. Hansen,  K. Jones.  2008.  Prioritizing avian conservation areas for the Yellowstone to Yukon Region of North America.  Biological Conservation 141: 908-924.Hansen, A.J. and R. DeFries. 2007. Land use change around nature reserves: Inplications for sustaining biodiversity. Ecological Applications, 17(4) 972-973.   PDF

    Hansen, A.J. and R. DeFries. 2007. Ecological mechanisms linking protected areas to surrounding lands. Ecological Applications 17(4), 974-988. PDF

    Hansen, A.J. and R. DeFries. 2007. Land use change around nature reserves: implications for sustaining biodiversity. 17(4), 972-973. PDF

    DeFries, R., A.J. Hansen, B.L. Turner, R.S. Reid and J. Liu. 2007.  Towards scientific principles for regional management of landscapes surrounding protected areas.  Ecological Applications 17(4) 1031-1038.    PDF

    Gude, P.H., A.J. Hansen, D.A. Jones.  2007.  Biodiversity consequences of alternative future land use scenarios in Greater Yellowstone.  Ecological Applications 17(4), 1004-1018.    PDF

    Powell, S.L., A.J. Hansen.  2007.  Conifer cover increase in the Greater Yellowstone Ecosystem: Frequency, rates, and spatial variation.  Ecosystems. 10:204-216.    PDF

    Brown, K., A.J. Hansen, R.E. Keane, L.J. Graumlich.  2006.  Complex interactions shaping aspen dynamics in the Greater Yellowstone Ecosystem.  Landscape Ecology.  Landscape Ecology 21:933–951.    PDF

    Gude, P.H., A.J. Hansen, R. Rasker, B. Maxwell. 2006. Rate and drivers of rural residential development in the Greater Yellowstone.  Landscape and Urban Planning 77:131-151.    PDF   

    Hansen, A.J., and D. Brown. 2005.  Land use change in rural America: An introduction. Ecological Applications 15(6):1849-1850.     PDF

    Hansen, A.J., R. Knight, J. Marzluff, S. Powell, K. Brown, P. Hernandez, and K. Jones. 2005. Effects of exurban development on biodiversity: Patterns, mechanisms, research needs. Ecological Applications 15(6): 1893-1905.    PDF

    Chornesky, E.A., A.M. Bartuska, G.H. Aplet, K.O. Britton, J. Cummings-Carlson, F.W. David, J.Eskow, D.R. Gordon, K. Gottschalk, R.A. Haack, A.J. Hansen, R.N. Mack, F.J. Rahel, M.A. Shannon, L.A. Wagner, and T.B. Wigley. 2005. Science Priorities for Reducing the Threat of Invasive Species to Sustainable Forestry. BioScience 55(4):335-348.     PDF

    DeFries, R., A.J. Hansen, A. Newton, M.C. Hansen. 2005. Isolation of protected areas in tropical forests over the last twenty years. Ecological Applications 15(1): 19-26.    PDF

    Wessels, K. J., R. S. De Fries, J. Dempewolf, L. O. Anderson, A. J. Hansen, S. L. Powell and E. F. Moran.  2004.  Mapping regional land cover with MODIS data for biological conservation: Examples from the Greater Yellowstone Ecosystem, USA and Pará State, Brazil.  Remote Sensing of Environment 92(1):67-83.     PDF

    Parmenter, A.P., A. Hansen, R. Kennedy, W. Cohen, U. Langner, R. Lawrence, B. Maxwell, A. Gallant, R. Aspinall. 2003. Land Use and Land Cover Change in the Greater Yellowstone Ecosystem: 1975-95. Ecological Applications 13(3):687-703.     PDF

    Gallant, A.L. A.J. Hansen, J.S. Councilman, D.K. Monte, and D.W. Betz. 2003. Vegetation dynamics under fire exclusion and logging in a Rocky Mountain watershed: 1856-1996. Ecological Applications 13(2):385-403.    PDF

    Hansen, A.J., R. Rasker, B. Maxwell, J.J. Rotella, J. Johnson, A. Wright Parmenter, U. Langner, W. Cohen, R. Lawrence, and M.V. Kraska. 2002. Ecological causes and consequences of demographic change in the New West. BioScience 52(2) 151-168.     PDF

    Hansen, A.J., and J.J. Rotella.  2002. Biophysical factors, land use, and species viability in and around nature reserves. Conservation Biology 16(4):1-12.     PDF

    Kramer, M.G., A.J. Hansen, M. Taper, and E. Kissinger. 2001.  Abiotic controls on windthrow and pattern and process of forest development on Kuiu Island, Alaska. Ecology 82(10):2749-2768.     PDF

    Hansen, A.J., and V. Dale. 2001. Biodiversity in U.S. Forests Under Global Climate Change. Ecosystems (4):161-163.     PDF

    Hansen, A.J., R.P. Neilson, V. Dale, C. Flather, L. Iverson, D. J. Currie, S. Shafer, R. Cook, P. Bartlein. 2001. Global change in forests: Interactions among biodiversity, climate, and land use. BioScience 51(9):765-779.     PDF

     Burrough, P.A., J.P. Wilson, P.F.M. van Gaans, A.J. Hansen. 2001. Fuzzy k-means classification of topo-climatic data as an aid to forest mapping in the Greater Yellowstone Area, USA. Landscape Ecology 16(6):523-546.     PDF

    Hansen, A.J., J.J. Rotella, M.L. Kraska and D. Brown. 2000. Spatial patterns of primary productivity in the Greater Yellowstone Ecosystem. Landscape Ecology. 15:505-522.      PDF

    Madden, E.M., R.K. Murphy, A.J. Hansen and L. Murray. 2000. Models for guiding management of prairie bird habitat in northwestern North Dakota. American Midland Naturalist 144:377-392.     PDF

    Rotella, J.J., M.L. Taper, and A.J. Hansen. 2000. Correcting nesting-success estimates for observer effects: maximum-likelihood estimates of daily survival rates with reduced bias. Auk 117:92-109. 

    Rasker, R. and A.J. Hansen. 2000. Natural amenities and population growth in the Greater Yellowstone region. Human Ecology Review 7(2):30-40.     PDF

    Hansen, A.J., J.J. Rotella, M.L. Kraska, and D. Brown. 1999. Dynamic habitat and population analysis: A filtering approach to resolve the biodiversity manager's dilemma. Ecological Applications 9(4):1459-1476.     PDF

    Madden, E.M., A.J. Hansen, and R.K. Murphy. 1999. Influence of prescribed fire history on habitat and abundance of passerine birds in northern mixed-grass prairie. Canadian Field Naturalist 113:627-640.     PDF

    Rotella, J.J., E.M. Madden, and A.J. Hansen. 1999. Sampling considerations for estimating abundance of passerines in grasslands. Studies in Avian Biology 19:237-243. 

    Hansen, A.J., R. Patten, and G. DeGayner, and B. Marks.  1996.  Simulating forest and habitat change in Southeast Alaska with the landscape model PAYSAGE.  Transactions in GIS 1(2):119-136.

    Hansen, A.J., W.C. McComb, R. Vega, M.G. Raphael, M. Hunter.  1995.  Bird habitat relationships in natural and managed forests in the west Cascades of Oregon.  Ecological Applications 5(3):555-569.    PDF

     Hansen, A.J., S.L. Garman, J.F. Weigand, D.L. Urban, W.C. McComb, and M.G. Raphael.  1995.   Ecological and economic effects of alternative silvicultural regimes in the PNW. Ecological Applications 5(3):535-554.

    Hansen, A.J.  1995.  Understanding and managing ecotones: Summary of the first SCOPE workshop on landscape boundaries.  Ecology International (22) 3-15.

    Hansen, A.J., S. Garman, B. Marks and P. Lee.  1993.  An approach for managing vertebrate diversity across multiple-use landscapes.  Ecological Applications. 3(3):481-496.

    Hansen, A.J, S.L. Garman, P. Lee and E. Horvath.  1993.  Do edge effects influence tree growth rates in Douglas-fir plantations?  Northwest Science 67(2):112-116.

    Hansen, A.J. and D.L. Urban. 1992.  Avian response to landscape pattern:  The role of species life histories.  Landscape Ecology 7:163-180.    PDF

    Hansen, A.J., T. Spies, F. Swanson and J. Ohmann.  1991.  Conserving biodiversity in managed forests.  BioScience 41(3):382-392.    PDF

    Hansen, A.J. et al. 1990. Responses of wildlife habitats to forest management and climate change: A modeling approach. Northwest Env. J. 6:419-420.

    Hansen, A.J., et al.  1990.  Responses of vertebrates to stand edge and type in managed forests of the Oregon Coast Range.  Northwest Env. J. 6:418-419.

    Hansen, A.J. 1987. Regulation of bald eagle reproductive rates in southeast Alaska.  Ecology 68:1387-1392.   PDF

    Hodges, J.E., E.L. Boeker and A.J. Hansen. 1987. Movements of radio- tagged bald eagles in and from southeast Alaska. Canadian Field Naturalist. 101:136-140.

    Hansen, A.J. 1986. Foraging behavior of bald eagles: A test of evolutionary game theory. Ecology 67:787-797.

    Hansen, A.J. and S. Rohwer. 1986. Coverable badges and resource defense in birds.  Animal Behavior 34:69-76.

    Hansen, A.J. 1986. Fighting behavior in Bald Ealges: A test of game theory. Ecology 67(3):787-797.   PDF

    Hansen, A.J. 1985. High rates of nonbreeding adult bald easgles in southeastern alaska.  Journal of Wildlife Management, 49(2):454-458.    PDF



    Hansen, A.J., and F. di Castri, editors.  1992.  Landscape Boundaries:  Consequences for Biotic Diversity and Ecological Flows.  Springer-Verlag Ecological Studies Series, New York. 432 pages. 

    di Castri, F., A.J. Hansen, M. Debussche, editors.  1990.  Biological Invasions in Europe and the Mediterranean Basin.  Kluwer Academic Publishers, Dordrecht, The Netherlands.



    Hansen, A. J. 2011. Contribution of source-sink theory to protected area science. Pgs 339-360 in J. Liu, V., Hull, A. Morzillo, and J. Wiens, eds. Sources, Sinks, and Sustainability across Landscapes. Cambridge University Press.

    Hansen, A.J.  2010.  Thinking big about Greater Yellowstone.  Pgs 1-16 in Knowing Yellowstone: Science in America’s First National Park. 2010 Jerry Johnson, Editor. Taylor Trade Publishing, Boulder, CO.

    Powell, S.L., A.J. Hansen, and W.B. Cohen.  2008.  Mapping the extent and distribution of conifer cover increase in the Greater Yellowstone Ecosystem.  In Landscape Ecology Research Trends.  Nova Science Publishers, Hauppauge, New York.  PDF

    Hansen, A.J., and R.S. DeFries.  2005.  Land use intensification around nature reserves in mountains: Implications for biodiversity.  Pgs. 551-562 in U. M. Huber, H.K.M. Bugmann, and M.A. Reasoner, eds., Global Change and Mountain Regions: An Overview of Current Knowledge. Kluwer Academic Publishers, New York.

    Hansen, A.J., R. DeFries, W. Turner. In Press. Land Use Change and Biodiversity: A Synthesis of Rates and Consequences during the Period of Satellite Imagery. Pgs xx-xx in G. Gutman and C. Justice, eds., Land Change Science: Observing, Monitoring, and Understanding Trajectories of Change on the Earth's Surface. Kluwer Academic Publishers, New York, NY.

    Hansen, A.J., and J.J. Rotella. 2002. Rural Development and Biodiversity: A Case Study from Greater Yellowstone. Pgs 123-140 in: Levitt, J., ed. Conservaton in the Internet Age. Island Press. New York, NY. 

    Hansen, A.J. and J.J. Rotella. 2001. Nature Reserves and Land Use: Implications of the "Place" Principle. Pages 57-75 in Dale, V. and R. Haeuber, eds. Applying Ecological Principles to Land Management. Springer Verlag, New York, NY.     PDF

    Joyce, L., J. Aber, S. McNulty, V. Dale, A. Hansen, L. Irland, R. Neilson, and K. Skog. 2001. Potential consequences of climate variability and change for the forests of the United States. Pgs 489-522 in National Assessment Synthesis Team. Climate Change Impacts on the United States: The Potential Consequences of Climate Variability and Change. Cambridge University Press, Cambridge, U.K. 

    Hansen, A.J. and J.J. Rotella. 2000. Bird responses to forest fragmentation. Pgs 202-221 in R.L. Knight, F.W. Smith, S.W. Buskirk, W.H. Romme, and W.L. Baker, (eds.) Forest Fragmentation in the Southern Rocky Mountains. University of Colorado Press, Boulder, CO.     PDF

    Hansen, A.J. and J.J. Rotella. 1999. Abiotic factors. Pages 161-209. In: M.L Hunter, Jr. (ed.). Maintaining biodiversity in forest ecosystems. Cambridge University Press, Cambridge, U.K. PDF

    Hansen, A.J., A. Gallant, J.J. Rotella, D. Brown. 1999. Natural and human drivers of biodiversity in the Greater Yellowstone Ecosystem. Pgs. 61-70 in Land use history of North America: Providing a context for understanding environmental change. T. Sisk, (ed.) Biological Resources Division, U.S. Geological Survey, Washington, D.C.