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BIOL
Biology
Department of Ecology
Department of Cell Biology and Neuroscience
Department of Plant Sciences & Plant Pathology




BIOL 100IN ORGANISM FUNCTION

BIOL 101IN ORGANISMAL BIOLOGY

BIOL 102   MOLECULAR & CELLULAR BIOLOGY

BIOL 103CS ENVIRONMENTAL SCIENCE & SOCIETY

BIOL 106CS INSECTS AND HUMAN SOCIETY (Previously ENTO 102CS)

BIOL 113   TWENTY FIRST CENTURY BIOLOGY

BIOL 195   INTEGRATIVE PHYSIOLOGY I

BIOL 204IN INSECT BIOLOGY

BIOL 207   ANATOMY & PHYSIOLOGY I

BIOL 208   ANATOMY & PHYSIOLOGY II

BIOL 213   INTRODUCTORY BIOLOGY:CELLS TO ORGANISMS

BIOL 214   INTRODUCTORY BIOLOGY:MOLECULES TO CELLS

BIOL 215   INTRODUCTORY BIOLOGY:ORGANISMS TO POPULATIONS

BIOL 230   IDENTIFICATION OF SEED PLANTS

BIOL 251   BOTANY: AN INTRODUCTION TO PLANT BIOLOGY

BIOL 252   BOTANY LAB

BIOL 280   SPECIAL TOPICS

BIOL 289R  UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY INSTRUCTION

BIOL 290R  UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY

BIOL 297   HUMAN PHYSIOLOGY I

BIOL 298   HUMAN PHYSIOLOGY II

BIOL 301   PRINCIPLES OF GENETICS

BIOL 302   ADVANCED CELL & MOLECULAR BIOLOGY

BIOL 303   PRINCIPLES OF ECOLOGY

BIOL 310   COMPARATIVE VERTEBRATE ANATOMY

BIOL 311   DEVELOPMENTAL BIOLOGY

BIOL 312   HISTOLOGY

BIOL 316RN INTRODUCTION TO RESEARCH IN MOLECULAR BIOLOGY 

BIOL 313   NEUROPHYSIOLOGY  (Offered previously as BIOL 413, effective Fall odd years.)                          

BIOL 395   HUMAN PATHOPHYSIOLOGY

BIOL 400   SEMINAR

BIOL 401   BIOLOGY INSTRUCTING

BIOL 403   EVOLUTION

BIOL 405   BEHAVIORAL & EVOLUTIONARY ECOLOGY

BIOL 406   ROCKY MOUNTAIN VEGETATION

BIOL 407   ALPINE ECOLOGY

BIOL 410   ADVANCED HUMAN ANATOMY

BIOL 411   ANIMAL PHYSIOLOGY

BIOL 413   NEUROPHYSIOLOGY

BIOL 415   ICHTHYOLOGY

BIOL 418   MAMMALOGY

BIOL 419   ORNITHOLOGY

BIOL 420   FIELD ORNITHOLOGY

BIOL 421   YELLOWSTONE WILDLIFE ECOLOGY

BIOL 422   GENES AND CANCER

BIOL 424   FRESHWATER ECOLOGY

BIOL 425   SENSORY NEUROPHYSIOLOGY

BIOL 426   NEUROETHOLOGY

BIOL 427   AQUATIC FIELD ECOLOGY

BIOL 430   PLANT PHYSIOLOGY

BIOL 435   INSECT IDENTIFICATION(Previously ENTO 432) 

BIOL 436   PLANT SYSTEMATICS

BIOL 437   PLANT DEVELOPMENT

BIOL 438   DEVELOPMENTAL MECHANISMS

BIOL 443   CURRENT TOPICS IN BIOLOGY

BIOL 445   COGNITIVE NEUROSCIENCE

BIOL 447   CONSERVATION BIOLOGY    

BIOL 448   CONSERVATION GENETICS    

BIOL 449   POPULATION GENETICS

BIOL 451   CELL BIOLOGY & NEUROSCIENCE DEPARTMENT CAPSTONE SEMINAR

BIOL 453R  BIOMIMETIC INTELLIGENT SYSTEMS

BIOL 455   PLANT ECOLOGY

BIOL 466R  GENE CONSTRUCTION (Effective Fall 2005)

BIOL 467   MOLECULAR MEDICINE

BIOL 470   INDEPENDENT STUDY

BIOL 480   SPECIAL TOPICS

BIOL 489R  UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY INSTRUCTION

BIOL 490R  UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY

BIOL 500   SEMINAR

BIOL 501   EVOLUTIONARY GENETICS

BIOL 502   ADVANCED LIMNOLOGY

BIOL 503   PALEOBIOLOGY

BIOL 504   QUANTITATIVE BIOLOGY

BIOL 505   ENVIRONMENTAL ANALYSIS

BIOL 506   POPULATION DYNAMICS

BIOL 507   COEVOLUTION

BIOL 508   SCIENTIFIC WRITING

BIOL 509   INTRODUCTION TO PRACTICAL MODELING

BIOL 510   TOPICS IN NEUROBIOLOGY

BIOL 513   TERRESTRIAL ECOLOGY OF PLAINS AND PRAIRIES

BIOL 515   LANDSCAPE ECOLOGY AND MANAGEMENT

BIOL 516   TERRESTRIAL ECOLOGY OF THE NORTHERN ROCKY MOUNTAINS

BIOL 518   PARAMETER ESTIMATION FOR ECOLOGICAL MODELS

BIOL 519   BIOLOGY OF RIPARIAN ZONES AND WETLANDS

BIOL 520   ANIMAL BIODIVERSITY IN GYE

BIOL 521   CONSERVATION BIOLOGY

BIOL 522   BIRDS OF PREY IN THE GREATER YELLOWSTONE ECOSYSTEM

BIOL 523   WILDLIFE ECOLOGY OF TE NORTHERN ROCKY MOUNTAINS

BIOL 524   FRONTIERS IN LANDSCAPE ECOLOGY

BIOL 525   RESEARCH METHODS AND THE SCIENTIFIC PROCESS

BIOL 532   PHYSIOLOGICAL PLANT ECOLOGY

BIOL 533   PHYSIOLOGICAL PLANT ECOLOGY LAB

BIOL 534   VEGETATION ECOLOGY

BIOL 540   ANALYSIS OF ECOLOGICAL COMMUNITIES

BIOL 542   COMMUNITY ECOLOGY

BIOL 548   CONSERVATION GENETICS

BIOL 570   INDEPENDENT STUDY

BIOL 575   RESEARCH OR PROFESSIONAL PAPER/PROJECT

BIOL 576   INTERNSHIP

BIOL 580   SPECIAL TOPICS

BIOL 588   PROFESSIONAL DEVELOPMENT

BIOL 589   GRADUATE CONSULTATION

BIOL 590   MASTER'S THESIS

BIOL 690   DOCTORAL THESIS


BIOL 100IN ORGANISM FUNCTION
F 3 cr. LEC 3
-- Comparison of plant and animal systems with respect to structure and function. The underlying chemistry, energetics, and ecological adaptations are discussed.

BIOL 101IN ORGANISMAL BIOLOGY
F,S 4 cr. LEC 3 LAB 1
-- Examination of five kingdoms of organisms (monera, protista, fungi, plants, animals), with concentrations on vascular plants and vertebrate animals. Considers survival strategies, nutrition, reproduction, and ecological and economic importance.

BIOL 102 MOLECULAR & CELLULAR BIOLOGY
F,S 4 cr. LEC 3 LAB 1
PREREQUISITE: CHMY 121 or CHMY 141.
-- Introduction to cellular organization and function. Topics covered include synthesis and function of macromolecules, cell organelles and structure, energy transformations in living systems, respiration, photosynthesis, the cell cycle, classical genetics, molecular genetics, and biotechnology.

BIOL 103CS ENVIRONMENTAL SCIENCE & SOCIETY
F,Su 3 cr. LEC 3
-- The relationship between people and the environment using the earth as an ecosystem to show the effects of people's activities on natural ecosystems. Environmental issues such as wilderness, wolf reintroduction, global warming, fire ecology, whirling disease, and grizzlies are covered.

BIOL 106CS INSECTS AND HUMAN SOCIETY (Previously ENTO 102CS)
S 3 cr. LEC 2 RCT 1
-- Ways in which research and advances in technology in the areas of insect biology and management have influenced people's lives throughout the world. Focus will be on insects as major factors affecting the areas of the world where humans live, crops and animals humans produce, and general quality of life on the planet. Interactions of insects and human cultures,technologically oriented and indigenous, non-technology based cultures, and concepts of pest management will also be explored. Students generate and test hypothesis and evaluate sources of scientific information on these topics.

BIOL 113 TWENTY FIRST CENTURY BIOLOGY
F 2 cr. LEC 2
-- Introductory freshman seminar designed to expose students to biology as a discipline, and establish the standards for scientific literacy. Students learn how hypotheses are formulated and tested, the nature and necessity for experimental controls, and what constitutes "burden of proof" in scientific inquiry.

BIOL 195 INTEGRATIVE PHYSIOLOGY I
F 4 cr. LEC 3 LAB 1
COREQUISITE: CHMY 141 or CHMY 151.
-- How the human body works. For students planning to be health professionals. Designed to be taken before Biol 213, includes basic cellular mechanisms, physiological control and communications. Major topics include muscle, nerve, respiratory, renal and cardiovascular systems. Cadaver laboratory will cover related human anatomy.

BIOL 204IN INSECT BIOLOGY
F 3 cr. LEC 2 LAB 1
PREREQUISITE: One of the following: BIOL 100, BIOL 101, or BIOL 102.
-- General biology of insects including principles of morphology, physiology, behavior, ecology, and control. Includes identification of major orders and common families.

BIOL 207 ANATOMY & PHYSIOLOGY I
S,Su 5 cr. LEC 3 LAB 2
PREREQUISITE: CHMY 121, CHMY 141 or CHMY 151 with a grade of "C-" or better. Priority given to majors requiring this course.
-- General principles of cell and tissue biology that apply to all living systems. Structure and function of skeletal, muscular, nervous, and endocrine systems. Homeostasis, control, and integration of the human body will be emphasized. Laboratory will cover related systems. This course is not repeatable without prior consent of instructor.

BIOL 208 ANATOMY & PHYSIOLOGY II
F 4 cr. LEC 3 LAB 1
PREREQUISITE: BIOL 102, BIOL 207, BIOL 214 or MB 301 with grades of "C-" or better in each course. Priority given to majors requiring this course.
-- Structure and function of digestive, cardiovascular, respiratory, reproductive, and urinary systems of humans. Principles of integration, metabolism, energy flow, and homeostasis will be emphasized. This course is not repeatable without prior consent of instructor.

BIOL 213 INTRODUCTORY BIOLOGY:CELLS TO ORGANISMS
S 4 cr. LEC 3 LAB 1
PREREQUISITE: STAT 216 and CHMY 141.
-- Introduction to form and function in the animal and plant kingdoms. Topics will include circulation and gas exchange, chemical signals, reproduction, nutrition, and the animal nervous system. Laboratories will be inquiry based with mathematical and computational applications to biological problems.

BIOL 214 INTRODUCTORY BIOLOGY:MOLECULES TO CELLS
F 4 cr. LEC 3 LAB 1
PREREQUISITE: STAT 216 and CHMY 141 and BIOL 213 and CHMY 143.
-- Introduction to biological macromolecules, cell structures and function, and gene structure and expression. The laboratory portion will include both wet labs and computer-based modules.

BIOL 215 INTRODUCTORY BIOLOGY:ORGANISMS TO POPULATIONS
S 4 cr. LEC 3 LAB 1
PREREQUISITE: BIOL 102 or BIOL 214, STAT 216 and M 161 or M 171.
-- An introductory course in ecology and evolution with in-depth coverage of topics in micro- and macroevolution, behavioral ecology, population ecology, community ecology, and biodiversity. The laboratory portion of the course will include material on biological diversity, evolutionary processes, and ecological processes.

BIOL 230 IDENTIFICATION OF SEED PLANTS
S 4 cr. LEC 2 LAB 2
PREREQUISITE: BIOL 101.
Identification of conifers, trees and shrubs, and herbaceous seed plants; determination by use of manuals; vocabulary, classification and nomenclature; and preparation and collection of seed plant specimens. Cross-listed with PS 250.

BIOL 251 BOTANY: AN INTRODUCTION TO PLANT BIOLOGY
F 3 cr. LEC 3
PREREQUISITE: BIOL 101N.
-- This course will provide a thorough overview of the fundamentals of plant and fungal biology from evolutionary, ecological, and physiological perspectives. Cross-listed with PS 251.

BIOL 252 BOTANY LAB
F 1 cr. LAB 1
COREQUISITE: BIOL 251.
-- This lab will feature plant ecology and also plants that have been used as model organisms in studies of basic biological processes. Cross-listed with PS 252.

BIOL 280 SPECIAL TOPICS
On Demand 1 - 4 cr. Maximum 12 cr.
PREREQUISITE: None required but some may be determined necessary by each offering department.
-- Courses not required in any curriculum for which there is a particular one-time need, or given on a trial basis to determine acceptability and demand before requesting a regular course number.

BIOL 289R UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY INSTRUCTION
F,S 1-3 cr. RCT may be repeated
-- Classroom instruction associated with directed undergraduate research/creative activity projects.

BIOL 290R UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY
F,S 1-6 cr. IND may be repeated
-- Directed undergraduate research/creative activity which may culminate in a written work or other creative project. Course will address responsible conduct of research.

BIOL 297 HUMAN PHYSIOLOGY I
S 3 cr. LEC 3
PREREQUISITE: CHMY 121 or CHMY 141 or CHMY 151 with grades of "C-" or better in either course.
--General principles of cell and tissue biology; function of skeletal, muscular, nervous, and endocrine systems. Homeostasis, control, and integration of the human body will be emphasized.

BIOL 298 HUMAN PHYSIOLOGY II
F 3 cr. LEC 3
PREREQUISITE: BIOL 102 or BIOL 207 or BIOL 214 or BIOL 297 or MB 301 with grades of "C-" or better in each course.
-- Function of the human digestive, cardiovascular, respiratory, reproductive, and urinary systems. Principles of integration, metabolism, energy flow and homeostasis will be emphasized.

BIOL 301 PRINCIPLES OF GENETICS
F,S 3 cr. LEC 3
PREREQUISITE: BIOL 102 or BIOL 214 or MB 301.
-- Introduction to classical and molecular genetics of eukaryotes, with emphasis on transmission genetics, the structure and regulation of genes, and mechanisms of genetic change.

BIOL 302 ADVANCED CELL & MOLECULAR BIOLOGY
S 3 cr. LEC 3
PREREQUISITE: BCHM 340 or consent of instructor.
-- In-depth study of cell structure and function.

BIOL 303 PRINCIPLES OF ECOLOGY
S 3 cr. LEC 3
PREREQUISITE: BIOL 101 or BIOL 215; M 121 or M 160 or M 170; Recommended: STAT 216 or PSPP 318.
-- Relation of organisms to their environment. The composition, structure, function and distribution of populations, communities, and ecosystems.

BIOL 310 COMPARATIVE VERTEBRATE ANATOMY
S 4 cr. LEC 2 LAB 2
PREREQUISITE: BIOL 101 or BIOL 215.
-- A comparative study of organ systems of vertebrates. Laboratory utilizes representative vertebrate types.

BIOL 311 DEVELOPMENTAL BIOLOGY
S 4 cr. LEC 4
PREREQUISITE: BIOL 101 or BIOL 215, and BIOL 102 or BIOL 214, and BIOL 301.
-- Developmental Biology: Introduction to the cell signaling pathways and morphogenetic processes that establish the basic vertebrate body plan. Includes hands-on study of chicken and frog embryos.

BIOL 312 HISTOLOGY
F 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 102 or BIOL 214, senior standing and consent of instructor.
-- Microscopic study of cells, tissues, and selected mammalian organs.

BIOL 313 NEUROPHYSIOLOGY F 3 cr. LEC 3
PREREQUISITE: BIOL 213 and BIOL 214.
-- Physiology of integrative mechanisms in nervous systems. Topics range from the mechanisms of synaptic transmission and action potential generation to the neural basic of learning and memory.

BIOL 316RN INTRODUCTION TO RESEARCH IN MOLECULAR BIOLOGY
S 3 cr. LEC 1 LAB 2
PREREQUISITE: BIOL 213 and BIOL 214 and CHMY 141 and CHMY 143.
-- Introduction to research methodologies and techniques used in modern biological science research. This laboratory course will focus on wet-lab techniques including DNA purification and analysis, gene library screening and PCR, and cover modes of computer based analysis.

BIOL 395 HUMAN PATHOPHYSIOLOGY
S 3 cr. LEC 2 IND 1 SEM 1
PREREQUISITE: BCHM 340 or consent of instructor.
-- Students will research two diseases of their own choosing and give a class presentation of their findings. The presentation normally includes diagnosis, pathophysiology, and treatment.

BIOL 400 SEMINAR
F,S 1 cr. SEM 1 Maximum 4 cr.
PREREQUISITE: Junior standing and as determined for each offering.
-- Topics offered at the upper division level which are not covered in regular courses. Students attend and discuss seminar presentations by professional biologists.

BIOL 401 BIOLOGY INSTRUCTING
F,S,Su 2 cr. LAB 2 Maximum 4 cr.
PREREQUISITE: Junior or senior standing, consent of instructor and department head.
-- Provides deeper contact with a subject for those considering an academic profession. This provides experience in a teaching laboratory under detailed academic supervision in recognition that teaching enhances learning. Includes the preparation, organization, presentation of materials, and student evaluation.

BIOL 403 EVOLUTION
S 3 cr. LEC 3
PREREQUISITE: BIOL 301.
-- For seniors in biology. Evolutionary theory is presented and takes two principle directions, the study of evolutionary history, and the study of natural selection.

BIOL 405 BEHAVIORAL & EVOLUTIONARY ECOLOGY
S 3 cr. LEC 3
PREREQUISITE: BIOL 303.
-- Abundance and distribution of organisms in relation to their evolution, behavior, population biology and interactions with other organisms.

BIOL 406 ROCKY MOUNTAIN VEGETATION
F 2 cr. LEC 1 LAB 1
PREREQUISITE: Junior or senior status in biological sciences and consent of instructor.
-- Field identification of major Rocky Mountain ecosystem-types; the composition, structure and function of climax and alternate communities; their environments, geography and history; and discussion of management alternatives. Includes introduction to field methods, statistical evaluations, remote sensing, and library use.

BIOL 407 ALPINE ECOLOGY
Su 3 cr. LEC 1 LAB 2
PREREQUISITE: Junior standing, BIOL 101.
-- The ecological characteristics of alpine areas. A three-day field trip will confirm and reinforce material presented in class and is a course requirement.

BIOL 410 ADVANCED HUMAN ANATOMY
S 4 cr. LEC 2 LAB 2.
PREREQUISITE: Senior standing, completion of at least two upper division courses in the biological sciences and consent of instructor.
-- Dissection and prosection of the detailed anatomy of the human thorax, abdomen and pelvis with special dissection emphasis on the extremities. Topographic, three-dimensional and detailed relationships of organs, nerves and vessels are emphasized. Can fulfill upper division honor credits.

BIOL 411 ANIMAL PHYSIOLOGY
F 3 cr. LEC 3
PREREQUISITE: BIOL 102 or BIOL 214, and one of the following: CHMY 211, CHMY 321, or CHMY 123.
-- General homeostatic physiology of animals with emphasis on mammals. Selected body systems are covered with major emphasis on the integration of body processes.

BIOL 413 NEUROPHYSIOLOGY F 3 cr. LEC 3 (To be offered as BIOL 313.)
PREREQUISITE: Junior standing and BIOL 411 or BIOL 207 or BIOL 214 or consent of instructor.
-- Physiology of integrative mechanisms in nervous systems. Topics range from the mechanisms of synaptic transmission and action potential generation to the neural basis of learning and memory.

BIOL 415 ICHTHYOLOGY
S 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 310.
-- Characteristics, classification, evolution, and life histories of major groups of marine and freshwater fishes, with an emphasis on North American freshwater fauna. Laboratory emphasizes identification, nomenclature, morphology, and distribution of Montana species.

BIOL 418 MAMMALOGY
F 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 310.
-- Evolution, functional biology, distribution, and classification of mammals. Labs cover taxonomy and identification of representative forms.

BIOL 419 ORNITHOLOGY
S 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 310.
-- Evolution, functional biology, distribution, and classification of birds. Montana species recognition is developed through laboratory use of a representative skin collection.

BIOL 420 FIELD ORNITHOLOGY
Su 3 cr. LAB 3
PREREQUISITE: Junior standing, and either BIOL 100 or BIOL 101.
-- Field identification, habitat affinities and life histories of birds of the northern Rockies. Includes early morning field trips.

BIOL 421 YELLOWSTONE WILDLIFE ECOLOGY
Su 3 cr. LEC 2 LAB 1
PREREQUISITE: Junior standing, and either BIOL 100 or BIOL 101.
-- Basic ecology of the major animal species of the Yellowstone area and the ecological controversies surrounding their management.

BIOL 422 GENES AND CANCER
F 3 cr. LEC 3
PREREQUISITE: BIOL 302.
-- This course will focus on the molecular and cellular mechanism of human cancer. The role of oncogenes and tumor suppressor genes in normal and cancerous cells will be examined, with an emphasis on how mutations in certain genes results in altered cell-cell signaling and cell proliferation. The role of genetic mutation in breast, colorectal and lymphoma cancers will be discussed, along with new technologies to detect and treat these cancers.

BIOL 424 FRESHWATER ECOLOGY
F 3 cr. LEC 3
PREREQUISITE: BIOL 303 or consent of instructor.
-- This course examines relationships between freshwater organisms and their environment. Students learn about the ecology of rivers, lakes, reservoirs, and wetlands, with exposure to a wide diversity of organisms and processes. Emphasis is placed on linking basic concepts and real-world applications.

BIOL 425 SENSORY NEUROPHYSIOLOGY
S 3 cr. LEC 3
PREREQUISITE: BIOL 313.
-- Neurophysiology of sensory cells and systems. Topics range from the mechanisms underlying sensory reception to the processing of sensory information at higher stages. The major focus will be on human sensory systems. Pathologies that effect sensory perception will be considered.

BIOL 426 NEUROETHOLOGY
On Demand 3 cr. LEC 3
PREREQUISITE: Consent of instructor.
-- Neural and hormonal bases of animal behavior, including mechanisms underlying sensory perception and motor responses, learning and memory, spatial navigation, language, dominance hierarchies and aggression, mating systems, and parental behavior. Model systems from a variety of different animal species will be studied including humans.

BIOL 427 AQUATIC FIELD ECOLOGY
F 2 cr. LEC 1 LAB 1
PREREQUISITE: Prior or concurrent registration in Biol 424.
-- Optional laboratory for BIOL 424. Introduction to representative freshwater habitats, communities, organisms, and sampling methods through laboratory and field exercises and classroom discussions. Formal written reports are required after completed exercises.

BIOL 430 PLANT PHYSIOLOGY
S 3 cr. LEC 3
PREREQUISITE: Junior standing, BIOL 101 and one of the following: CHMY 211, CHMY 321, or CHMY 123.
-- Physiological processes of higher plants, including photosynthesis, water relations, mineral nutrition, and development. Cross-listed with PS 450.

BIOL 435 INSECT IDENTIFICATION(Previously ENTO 432)
S alternate years, to be offered odd years 4 cr. LEC 2 LAB 2
PREREQUISITE: ENTO 204N and one of the following: BIOL 100, BIOL 101, or BIOL 102.
-- The identification of insects and related terrestrial arthropods. Evolutionary patterns reflected in modern insect diversity will be used to illustrate classification methods. Taxonomic methods will be used as an access to information retrieval.

BIOL 436 PLANT SYSTEMATICS
F alternate years, to be offered even years 3 cr. LEC 1 LAB 2
PREREQUISITE: BIOL 101 and BIOL 230.
-- Introduction to the local vascular plant flora emphasizing characteristics of the common families and genera. Lab concentrates on plant identification of common angiosperm plant families in Montana; preparation of about 120 reference specimens taken from the local flora. Cross-listed with PS 456.

BIOL 437 PLANT DEVELOPMENT
F alternate years, to be offered even years 3 cr. LEC 3
PREREQUISITE: BIOL 301.
-- Cellular and molecular mechanisms of the development of multi cellular life forms that consist of walled cells, and primarily plants. Some topics include developmental differences between plants and animals, regulation of gene expression, environmental effects on plant development, and computer modeling of development. Cross-listed with PS 457.

BIOL 438 DEVELOPMENTAL MECHANISMS
F 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 302.
-- This course will focus on the molecular and cellular mechanisms which drive developmental processes.

BIOL 443 CURRENT TOPICS IN BIOLOGY
S 2 cr. SEM 2
PREREQUISITE: Senior standing in Ecology Department, and prior or concurrent registration in BIOL 403.
-- Senior capstone course. Discussion of topics that integrate evolutionary theory with ecology, genetics, medicine, behavior, or other subjects that are part of the biology curriculum.

BIOL 445 COGNITIVE NEUROSCIENCE
S 3 cr. LEC 3
PREREQUISITE: BCHM 340 plus one of: BIOL 208, PSYX 350, BIOL 411.
-- This course will survey our present knowledge of the neural basis of normal and abnormal cognitive function in humans and non-human primates. Topics will range from perception and action to attention, consciousness and mental illness.

BIOL 447 CONSERVATION BIOLOGY
F 3 cr. LEC 3
PREREQUISITE: BIOL 303.
-- Examines issues relevant to conservation of wild populations, focusing primarily on animals. Emphasis is on approaches that use demography, population biology and genetics to address conservation questions. Readings are from the primary literature, rather than a textbook, including case studies. Cross-listed with BIOL 521.

BIOL 448 CONSERVATION GENETICS
F 3 cr. LEC 3
PREREQUISITE: BIOL 301 and STAT 216.
-- Introduces the theory and practice of conservation genetics, focusing primarily on animals. Case studies will be used liberally, and emphasis will be placed on interpreting genetic data. Readings will include primary literature.

BIOL 449 POPULATION GENETICS
F alternate years, to be offered even years 3 cr. LEC 3
PREREQUISITE: BIOL 301.
-- Introduction to theory and empirical data on genetics of populations. Topics covered include modeling natural and artificial selection, nonrandom mating, gene flow and effective population size as factors influencing the maintenance of genetic variation in populations. The approach emphasizes the development of simple mathematical models to illustrate fundamental conceptual issues in the field.

BIOL 451 CELL BIOLOGY & NEUROSCIENCE DEPARTMENT CAPSTONE SEMINAR
F,S 2 cr. SEM 2
PREREQUISITE: Senior standing in the Cell Biology & Neuroscience Department.
-- Senior capstone course. Students are expected both to present and to discuss advanced topics from the current biomedical literature. These topics will expand upon material presented in regular courses in the biomedical science curriculum. Students will write at least one major paper.

BIOL 453R BIOMIMETIC INTELLIGENT SYSTEMS
F 4 cr. LEC 1 LAB 3
PREREQUISITE: CAPP 120, M 170 or M 176 or M 182, STAT 216 or STAT 332.
COREQUISITE: BIOL 313 or CS 436 or EE 308.
-- Students will study behaviors, structures and organs of simple biological organisms and model key aspects of these organisms in biomimetic robots. Students will gain hands-on experience with mathematical, engineering and software tools, all in the context of biological modeling.

BIOL 455 PLANT ECOLOGY
S 3 cr. LEC 3
PREREQUISITE: BIOL 101 or BIOL 215 and BIOL 303 or ARNR 240.
-- Principles of plant ecology, covering plant-environment relations, plant life histories, plant species interactions, plant community concepts, succession, and the role of plants in ecosystem processes.

BIOL 466R GENE CONSTRUCTION
F 3 cr. LAB 3
PREREQUISITE: BIOL 302 or BCHM 340.
-- The goals are to provide upper level students with the opportunity of designing and building their own genes. The goal of the course is to use this design experience to learn basic techniques in cell and molecular biology.

BIOL 467 MOLECULAR MEDICINE
S 3 cr. LEC 1 SEM 2
PREREQUISITE: BIOL 301 and BCHM 340.
-- Lecture and seminar courses based on recent, original papers. Moves from human disease to molecular explanations. Intended for upper level students with a strong background in biology.

BIOL 470 INDEPENDENT STUDY
On Demand 1 - 3 cr. IND Maximum 6 cr.
PREREQUISITE: Junior standing, consent of instructor and approval of department head.
-- Directed research and study on an individual basis.

BIOL 480 SPECIAL TOPICS
On Demand 1 - 4 cr. Maximum 12 cr.
PREREQUISITE: Course prerequisites as determined for each offering.
-- Courses not required in any curriculum for which there is a particular one-time need, or given on a trial basis to determine acceptability and demand.

BIOL 489R UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY INSTRUCTION
F,S,Su 1 - 2 cr. RCT May be repeated. Max 4 cr.
-- Classroom instruction associated with directed undergraduate research/creative activity projects.

BIOL 490R UNDERGRADUATE RESEARCH/CREATIVE ACTIVITY
F,S,Su 1 - 6 cr. IND May be repeated. Max 12 cr.
-- Directed undergraduate research/creative activity which may culminate in a research paper, journal article, or undergraduate thesis. Course will address responsible conduct of research.

BIOL 500 SEMINAR
On Demand 1 cr. SEM Maximum 4 cr.
PREREQUISITE: Graduate standing or seniors by petition and course prerequisites as determined for each offering.
-- Topics offered at the graduate level which are not covered in regular courses. Students participate in preparing and presenting discussion material.

BIOL 501 EVOLUTIONARY GENETICS
F alternate years, to be offered even years 3 cr. LEC 3
PREREQUISITE: BIOL 301.
-- The operations of fundamental genetic principles in populations of living things. Emphasis on natural populations with a thorough consideration of factors of evolution, particularly at the species level and below.

BIOL 502 ADVANCED LIMNOLOGY
S alternate years, to be offered even years 4 cr. LEC 2 LAB 2
PREREQUISITE: M 170, BIOL 404, BIOL 427, CHMY 123 or BCHM 340.
-- Advanced quantitative study of the physical, chemical and biological dynamics of lakes and reservoirs.

BIOL 503 PALEOBIOLOGY
S alternate years, to be offered odd years 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 310, BIOL 403.
-- A study of the fossil record as a means of inferring biological characteristics of extinct species. Current topics in paleontology, phylogenetic systematics, patterns of evolution, speciation and extinction and osteohistology will be examined.

BIOL 504 QUANTITATIVE BIOLOGY
F alternate years, to be offered even years 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 303, either STAT 216 or STAT 332, and one of the following: M 161, M 191, M 192.
-- Applications of mathematical models to biological phenomena with examples drawn from physiology, ecology and bioengineering. The course is intended to develop facility with optimization techniques, numerical methods, matrix operations, complex variables and simple statistical ideas. Computer lab.

BIOL 505 ENVIRONMENTAL ANALYSIS
S alternate years, to be offered even years 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 303, either STAT 216 or STAT 332 and one of the following: M 161, M 191, M 192.
-- Theory of ecological dynamics and introduction to multivariate methods for ecological analysis. Computer lab.

BIOL 506 POPULATION DYNAMICS
S alternate years, to be offered odd years 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 303, either STAT 216 or STAT 332, and one of the following: M 161, M 171, M 172.
-- Techniques for modeling the growth, regulation, harvesting and persistence of populations. Computer lab.

BIOL 507 COEVOLUTION
F alternate years, to be offered even years 3 cr. LEC 3
PREREQUISITE: BIOL 301, BIOL 303, or BIOL 403.
-- Exploration of nature and dynamics of symbiotic relationships between and among plants, animals, fungi and bacteria.

BIOL 508 SCIENTIFIC WRITING
S 3 cr. LEC 2 LAB 1
PREREQUISITE: Graduate standing and permission of instructor.
-- The course is designed to provide an intensive scientific writing experience for graduate students in the process of writing the first draft of their thesis or dissertation, including editing and critique sessions by both peers and the instructor.

BIOL 509 INTRODUCTION TO PRACTICAL MODELING
F 3 cr. LEC 2 LAB 1
PREREQUISITE: First courses in calculus and statistics or consent of instructor.
-- With computers, the power of mathematical modeling is accessible to every biologist. We will discuss philosophies, strategies, techniques and pitfalls of modeling. After this course, students should be able to answer complex biological questions by formulating and analyzing mathematical/computational models.

BIOL 510 TOPICS IN NEUROBIOLOGY
S 3 cr. LEC 2 RCT 1 Maximum of 9 credits.
PREREQUISITE: Graduate standing and at least one upper division or graduate course in neurobiology.
-- Recent advances in topics in neurobiology with emphasis in different years on either neurocytology, neuroendocrinology/neuroimmunology, or developmental neurobiology.

BIOL 513 TERRESTRIAL ECOLOGY OF PLAINS AND PRAIRIES
Su 1 cr. RCT 1
PREREQUISITE: Either BIOL 406 or BIOL 516, graduate standing, secondary teacher certification, two years teaching experience, and computer access.
COREQUISITE: Suggested: ESCI 513.
-- Students will develop plant keys for classroom use, quantitatively analyze two grassland communities, and develop classroom activities on ecology of grasslands. Distance learning, class offered by internet connection. This course is designed for secondary school teachers enrolled in MSSE program and cannot be used in graduate programs in Biological Sciences.

BIOL 515 LANDSCAPE ECOLOGY AND MANAGEMENT
F alternate years, to be offered odd years 4 cr. LEC 2 LAB 2
PREREQUISITE: Graduate standing or consent of instructor.
-- Principles on landscape pattern, change, and function. Application of theory to conservation including population viability, reserve design, multiple-use landscapes. Lab introduces GIS, GPS, and simulation models. For graduate students and motivated undergraduates.

BIOL 516 TERRESTRIAL ECOLOGY OF THE NORTHERN ROCKY MOUNTAINS
Su 2 cr. RCT 1 LAB 1
PREREQUISITE: Graduate standing, two years of classroom teaching, undergraduate science degree, and one year of biology.
-- Description and comparison of grass land, forest, and alpine ecosystems of the NRM with respect to composition, structure, and process such as production, decomposition and mineral cycling. We will use tools including keys to species and environmental types, dimension analysis, remote sensing, and statistics. This course is designed for secondary school teachers enrolled in MSSE Program and cannot be used in graduate programs in biological sciences.

BIOL 518 PARAMETER ESTIMATION FOR ECOLOGICAL MODELS
F alternate years, to be offered odd years 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 504 or equivalent.
-- Statistical methods to quantify uncertainty, and to plan data collection for cost-efficient reduction in uncertainty, in application to ecological models where data are often sparse and processes are often noisy, and management decision must take account of uncertainty.

BIOL 519 BIOLOGY OF RIPARIAN ZONES AND WETLANDS
Su 2 cr. RCT 2
PREREQUISITE: Either BIOL 516 or BIOL 406, secondary teacher certification, two years teaching experience, and computer access.
COREQUISITE: Suggested: ESCI 512, ESCI 515.
-- Students will develop plant keys for classroom use, quantitatively analyze two riparian and two wetland areas, and develop classroom activities about ecology of those areas. Distance learning class offered by internet connection. This course is designed for secondary school teachers enrolled in the MSSE program and cannot be used in graduate programs in Biological Sciences.

BIOL 520 ANIMAL BIODIVERSITY IN GYE
Su 2 cr. LEC 1 LAB 1
PREREQUISITE: BIOL 303, F&WL 301, BIOL 405, or equivalent and (a) 2 years science technology experience or (b) enrolled in MSSE.
-- Exploration of biodiversity's meaning, importance & determinants; key ecological features of the Greater Yellowstone Ecosystem and patterns of change in those features: & possible strategies for maintaining biodiversity in the Greater Yellowstone Ecosystem.

BIOL 521 CONSERVATION BIOLOGY
F 3 cr. LEC 3
PREREQUISITE: BIOL 303, BIOL 403 and STAT 216, or equivalents.
-- A broad survey of conservative biology, with studies spanning genetics, demography/population dynamics, and community/ecosystem/landscape ecology. Approaches include empirical field studies, mathematical models and conceptual discussion. Includes lab modeling exercises, extensive reading in primary literature, and writing a research paper. Cross-listed with BIOL 447.

BIOL 522 BIRDS OF PREY IN THE GREATER YELLOWSTONE ECOSYSTEM
Su 2 cr. LEC 1 LAB 1
PREREQUISITE: BIOL 303, FWL 301, BIOL 405, or equivalent and 2 years science technology experience or enrolled in MSSE.
-- Exploration of the ecology and habitat of avian raptors in the Greater Yellowstone Ecosystem (GYE). Application of the scientific method to the study of raptors. Field identification of raptors, investigation of species life histories, and inquiry methods of species-specific habitat needs. Student will develop methods and skills for classroom based research on wildlife. This course is designed for secondary school teachers enrolled in the MSSE program.

BIOL 523 WILDLIFE ECOLOGY OF THE NORTHERN ROCKY PLAINS
Su 2 cr. LEC 2
PREREQUISITE: BIOL 303, FWL 301, BIOL 405, or equivalent and 2 years science technology experience or enrolled in MSSE.
-- Introduction to wildlife species and the range of habitats present in the Northern Rocky Mountain ecosystems. Emphasis on large carnivores and ungulates within montane terrestrial systems. Application of the scientific method to study interactions between predators, prey , and human impacts. This course is designed for middle and high school teachers.

BIOL 524 FRONTIERS IN LANDSCAPE ECOLOGY
F alternate years to be offered even years 3 cr. LEC 2 LAB 1
PREREQUISITE: BIOL 303 or the equivalent.
-- Students and instructor will write a scientific paper for publication that synthesizes an important question in landscape ecology. Students will select the topic, review and synthesize current knowledge on the topic, and write a scientific manuscript.

BIOL 525 RESEARCH METHODS AND THE SCIENTIFIC PROCESS
F 3 cr. LEC 2 RCT 1
PREREQUISITE: Graduate standing or consent of instructor.
-- Application of the scientific method to answer biological questions and the development of skills needed to prepare research proposals, critique research studies and communicate research findings. For first year graduate students.

BIOL 532 PHYSIOLOGICAL PLANT ECOLOGY
F alternate years, to be offered odd years 3 cr. LEC 3
PREREQUISITE: BIOL 303.
-- Outlines the plant's Hutchinsonian niche through review of energy, material (water, nutrients and toxins) and mechanical (including animal) factors. Computer modeling of plant function in the environment is discussed.

BIOL 533 PHYSIOLOGICAL PLANT ECOLOGY LAB
F alternate years, to be offered odd years 1 cr. LAB 1
COREQUISITE: BIOL 532.
-- A research project in physiological plant ecology will be chosen, carried out and reported in scientific journal format.

BIOL 534 VEGETATION ECOLOGY
F alternate years, to be offered even years 3 cr. LEC 3
PREREQUISITE: BIOL 303.
-- Considers the composition, structure, function, distribution in time and space, ecology and classification of communities. Emphasizes universal methods, current studies and Rocky Mountain systems. Complementary field experience is available in BIOL 406.

BIOL 540 ANALYSIS OF ECOLOGICAL COMMUNITIES
F alternate years, 3 cr. LEC 2 LAB 1
-- Multivariate statistical analysis of data from terrestrial or aquatic, plant or animal communities. Classification, ordination, and predictive modeling of species and communities, emphasizing a hands-on approach and practical problem solving in community ecology.

BIOL 542
S alternate years, to be offered odd years 3 cr. LEC 3
PREREQUISITE: At least one undergraduate or graduate course in each of the following: ecology (e.g., M 170) and statistics (e.g. STAT 216)or consent of instructor.
--Focuses on the origin, maintenance, and consequences of biological diversity within local communities by examining studies of natural patterns, explorations of mathematical models and direct experimentation. The complexities of species interactions are explored in multispecies assemblages.

BIOL 548 CONSERVATION GENETICS
F 3 cr. LEC 3
PREREQUISITE: BIOL 301, STAT 216 and STAT 217.
-- Introduction to the application of genetics for the conservation of plant and animal populations. Emphasis will be placed on case studies from the primary literature and analyzing genetic data using mathematical models developed in class.

BIOL 570 INDEPENDENT STUDY
On Demand 1 - 3 cr. IND Maximum 6 cr.
PREREQUISITE: Graduate standing, consent of instructor, approval of department head and Dean of Graduate Studies.
-- Directed research and study on an individual basis.

BIOL 575 RESEARCH OR PROFESSIONAL PAPER/PROJECT
F,S,Su 1 - 4 cr. IND
Graduate standing and committee approval.
-- A research or professional paper or project dealing with a topic in the field. The topic must be mutually agreed upon by the student and his or her major advisor and graduate committee.

BIOL 576 INTERNSHIP
On Demand 2 - 12 cr. IND
PREREQUISITE: Graduate standing, consent of instructor and approval of department head.
-- An individualized assignment arranged with an agency, business or other organization to provide guided experience in the field.

BIOL 580 SPECIAL TOPICS
On Demand 1 - 4 cr. Max 12 cr.
PREREQUISITE: Upper division courses and others as determined for each offering.
-- Courses not required in any curriculum for which there is a particular one time need, or given on a trial basis to determine acceptability and demand before requesting a regular course number.

BIOL 588 PROFESSIONAL DEVELOPMENT
On Demand 1 - 3 cr. May be repeated; maximum 3 cr.
PREREQUISITE: Graduate standing, teaching experience and/or current employment in a school organization, consent of instructor and Dean of Graduate Studies.
-- Courses offered on a one time basis to fulfill professional development needs of in service educators. A specific focus is given to each course which is appropriately subtitled.

BIOL 589 GRADUATE CONSULTATION
F,S,Su 3 cr. IND
PREREQUISITE: Master's standing and approval of the Dean of Graduate Studies.
-- This course may be used only by students who have completed all of their course work (and thesis, if on a thesis plan) but who need additional faculty or staff time or help.

BIOL 590 MASTER'S THESIS
F,S,Su 1 - 10 cr. IND Maximum credits unlimited.
PREREQUISITE: Master's standing.

BIOL 690 DOCTORAL THESIS
F,S,Su 1 - 10 cr. IND Maximum credits unlimited.
PREREQUISITE: Doctoral standing.

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