Email: [email protected]


  • BA, Occidental College, 1974; Biology and German
  • Ph.D., University of Washington, Department of Biological Structure, 1979; Anatomy and Neuroscience                                                         
  • Postdoctoral in Neuroscience, Vanderbilt University in the laboratory of Jon Kaas, 1979-1983                                                         
  • Research Assistant Professor, Department of Psychology, Vanderbilt University, 1983-1985                  
  • Assistant Professor, Department of Anatomy, Tulane University School of Medicine, 1985-1987                  
  • Associate Professor, Department of Anatomy (Structural  and Cellular Biology), Tulane University School of Medicine, 1985-1993                                                         
  • Professor, Department of Anatomy, Tulane University School of Medicine, 1993-2008
  • Professor, Department of Cell Biology and Neuroscience, Montana State University, 2008- Present